CLASTOPUS Bunge ex Boiss., Fl. Orient. 1: 261 (1867).

Herbs perennial, cespitose, pulvinate, with few- to several-branched woody caudex covered with leaves or their remains of previous seasons. Trichomes subsessile or sessile, appressed, rigid, 8–12-rayed stellate, those on fruit with longer stalks and much softer, some mainly along proximal half of leaf margin, pedicels, and sepals appear simple but they represent modified stellate trichomes with one or rarely two rays much longer and the rest become rudimentary. Multicellular glands absen. Stems erect to ascending, simple, leafy, not spiny. Basal leaves short petiolate, rosulate, simple, entire; cauline leaves sessile, cuneate to attenuate, not auriculate at base, entire. Racemes several to many flowered, ebracteate, corymbose, elongated in fruit; rachis straight, pubescent; fruiting pedicels, ascending to divaricate, straight, pubescent all around, persistent. Sepals oblong, free, persistent well after fruit dehiscence, slightly to strongly accrescent and becoming lanceolate in fruit, distinctly larger than fruit, unequal, base of lateral pair slightly saccate; petals yellow or purple, erect at base with flaring blade, longer than sepals; blade oblanceolate, apex obtuse; claw equaling or slightly longer than sepals, glabrous, unappendaged, entire; stamens 6, slightly exserted, erect, tetradynamous; filaments winged to middle or upper third, toothed at end of winge, glabrous, free; anthers oblong, not apiculate; nectar glands 4, lateral, 1 on each side of lateral stamens, median glands absent; ovules 4&ndash8 per ovary; placentation parietal. Fruit indehiscent or tardily dehiscent, capsular silicles, ellipsoid to ovoid, terete or slightly latiseptate, not inflated, unsegmented; valves thickened, subleathery, veinless, glabrous or pubescent outside, glabrous inside, not keeled, apex acute and often extending in stylar area, smooth, wingless, unappendaged; gynophore absent; replum rounded, concealed by connate valve margins; septum complete, membranous, veinless; style 2–7 mm long, filiform, persistent, pubescent throughout to at least proximal third; stigma strongly 2-lobed, the lobes connivent, free, not or decurrent, opposite replum, unappendaged. Seeds biseriate, wingless, ovoid, plump; seed coat rugose, verrucose, not mucilaginous when wetted; cotyledons accumbent.